Scientia Marina, Forthcoming Articles

Distribution and spatio-temporal biomass trends of red mullets across the Mediterranean

George Tserpes
Hellenic Centre for Marine Research, Institute of Marine Biological Resources and Inland Waters, Greece

Enric Massutí
Instituto Español de Oceanografía, Centre Oceanogràfic de les Balears, Spain

Fabio Fiorentino
Italian National Research Council (CNR), Institute for Coastal Marine Environment (IAMC), Italy

Maria Teresa Facchini
COISPA Tecnologia & Ricerca, Italy

Claudio Viva
Centro Interuniversitario di Biologia Marina ed ecologia applicata (CIBM), Italy

Angélique Jadaud
IFREMER-UMR MARBEC, Fisheries Laboratory, France

Aleksandar Joksimovic
University of Montenegro-Institute of Marine Biology, Montenegro

Paola Pesci
Department of Life and Environmental Sciences, University of Cagliari, Italy

Corrado Piccinetti
Laboratorio di Biologia Marina e Pesca, BiGEA, Università di Bologna, Italy

Letizia Sion
Department of Biology, University of Bari Aldo Moro, Italy

Ioannis Thasitis
Department of Fisheries and Marine Research, Cyprus

Nedo Vrgoc
Institute of Oceanography and Fisheries, Croatia


The present work examines the spatio-temporal biomass trends of Mullus barbatus and Mullus surmuletus in the Mediterranean Sea through the analysis of a time series of data coming from the Mediterranean International Trawl Surveys (MEDITS), accomplished annually from 1994 to 2015. The biomass of both species showed clear declining trends below 150 to 200 m depth, which were steeper in the case of M. barbatus. Increases in temporal biomass trends were observed for M. barbatus from 2008 onward in most geographic sub-areas (GSAs), while stability was mostly observed for M. surmuletus. For both species, dynamic factor analysis revealed similarities among neighbouring GSAs and the subsequent cluster analysis identified two major GSA groups corresponding to the eastern and western basins of the Mediterranean. Overall, the results suggested that the combined effects of fishing and environmental conditions determine species abundance variations, but the relative importance of each component may vary among areas.


red mullet; striped red mullet; distribution; trends; Mediterranean

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